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Specimen Type: holotype Institution: BMNH British Museum Natural History, London, UK Photographer: Larry G. Bezark Collection Location: Ellopura, Sabah, Borneo 47525
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Institution: Heffern Collection Photographer: Dan Heffern Collection Location: Sabah, Borneo 71755
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Institution: LGB Larry G. Bezark Collection, Sacramento, California, USA Collection Location: Sabah, Borneo, Malaysia 36255
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Photographer: Ulf Nylander 36256
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